The ruins of Anuradhapura are one of South Asia’s most evocative sights. The sprawling complex contains a rich collection of archaeological and architectural wonders: enormous dagobas (brick stupas), ancient pools and crumbling temples, built during Anuradhapura’s thousand years of rule over Sri Lanka. Today, several of the sites remain in use as holy places and temples; frequent ceremonies give Anuradhapura a vibrancy that’s a sharp contrast to the museum-like ambience at Polonnaruwa.
Dambulla’s famed rock cave temple is an iconic Sri Lankan image – you’ll be familiar with its spectacular Buddha-filled interior long before you arrive in town. Despite its slightly commercial air, this remains an important holy place and should not be missed.
Much less visited than nearby Yala National Park, Bundala National Park is an excellent choice for birders, and you've a good chance of spotting crocs, wild boar, mongooses, monitor lizards, monkeys and elephants. Most people visit on jeep tours from Tissamaharama. Bundala is open year-round, allowing wildlife junkies to get a wet-season fix.
Weligama (meaning ‘Sandy Village’) is an interesting blend of lively fishing town and beach resort. The sprawling main settlement and coastal road is somewhat scruffy and not that easy on the eye, but you'll find the sandy beach is attractive once you're away from the main section; there's a couple of cove beaches west of the centre. At the east end of the beach, there's an enormous new concrete Marriott hotel, which towers over the surrounding buildings and looks completely out of place.
Rising dramatically from the central plains, the enigmatic rocky outcrop of Sigiriya is perhaps Sri Lanka's single most dramatic sight. Near-vertical walls soar to a flat-topped summit that contains the ruins of an ancient civilization, thought to once have been the epicenter of the short-lived kingdom of Kassapa, and there are spellbinding vistas across mist-wrapped forests in the early morning.
Often referred to as ‘Little England’, this genteel highland community does have a rose-tinted, vaguely British-country-village feel to it, with its colonial-era bungalows, Tudor-style hotels, well-tended hedgerows and pretty gardens. Indeed, Nuwara Eliya was once was the favored cool-climate escape for the hard-working and hard-drinking English and Scottish pioneers of Sri Lanka’s tea industry.
Trincomalee (Trinco) sits on one of the world’s finest natural harbors. This historic city is old almost beyond reckoning: it’s possibly the site of historic Gokana in the Mahavamsa (Great Chronicle), and its Shiva temple the site of Trikuta Hill in the Hindu text Vayu Purana. It makes a great stop over on the way to the nearby beaches of Uppuveli and Nilaveli.
In Tissamaharama (usually shortened to Tissa), eyes are automatically drawn upwards and outwards. Upwards to the tip of its huge, snowy-white dagoba and outwards, beyond the town’s confines, to nearby wildlife reserves crawling with creatures large and small. With its pretty lakeside location, Tissa is an ideal mellow base for the nearby Yala and Bundala National Parks.
Although it's unlikely it will reclaim its 19th-century moniker 'the garden city of the East', Colombo has nevertheless emerged as a must-see stop in Sri Lanka. No longer just the sprawling city you have to endure on your way to the beaches, it has become a worthy destination in its own right and makes an excellent start – or finish – to your Sri Lankan adventures.
Historic Batticaloa, Batti for short, enjoys a spectacular position surrounded by lagoons with palm-filtered sunlight glancing off the water. There's a mellow vibe to the town, and the compact center and its huge fortress and many churches are well worth a half-day's exploration on foot.